For the first time for the Baikal Lake basin, evidence of the existence of populations of sculpin Cottus cf. poecilopus Heckel (alpine bullhead) characteristic of the adjacent Lena River basin in large northern Baikal tributaries, the Verkhnaya (Upper) Angara and the Kichera is provided. In June 2009 during the study of the lower reaches of the Kholodnaya River (Kichera-Baikal system), 8 individuals of sculpin aged from 3 to 5 years and in August 2009 5 more specimens aged from 4 to 6 were caught. All fish were fertile with the gonads in maturity stage III. In August 2010 33 individuals aged from 4 to 6 were collected in the main channel of the Upper Angara near Novy Uoyan settl.. At about the same time C.poecilopus was registered in stomachs of Arctic charr from Lake Amut (Churo-Upper Angara-Baikal system) near the divide with the Pravaya (Right) Mama (Mama-Vitim-Lena system). In August 2010 17 specimens aged from 1 to 4 were caught in the Upper Angara near the mouth of its large left tributary the Yanchui River. Sculpins from the rivers Kholodnaya, Upper Angara and Yanchui have higher growth rate as compared to the ones from mountain lakes of the upper Lena basin. About 90 % of males and females matured at the age of 3 years. Absolute fecundity of two females from the Upper Angara River was 149 and 556 (with a mean of 352,5) eggs and of two females from the Kholodnaya River, 223 and 305 (with a mean of 264) eggs. This exceeds the fecundity of sculpins from mountain lakes in the Lena part of Baikal rift zone, which averages less than 150 eggs. Sculpins spawn in the Kholodnaya River in the 1st half of June. The diet of C. cf. poecilopus all over the Baikal basin as well as in adjacent sites in the Lena basin was basically composed of larvae of amphibious insects (trichopterans, ephemeropterans, chironomids and plecopterans). The discovery of the third species from the Lena basin in the Baikal basin following the findings of Arctic charr Salvelinus alpinus and grayling Thymallus baicalolenensis evidences the absence of differences in the structure of ichthyofaunas of the upper parts of both neighboring basins. Sculpins permeate along the streams to mountain lakes more easily than other species and colonize the most elevated ones in lake cascades within the northern part of Baikal rift zone. Their dispersal across the divide could proceed in two ways: via headwater captures or via flattened passes between converging upper reaches of adjacent streams.

Andreev Rostislav Sergeevich, Leading Engineer, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: andreev_rs@mail.ru 

Matveev Arkadiy Nikolaevich, Doctor of Science (Biology), Professor, Head of Chair, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: matvbaikal@mail.ru 

Samusenok Vitaliy Petrovich, Candidate of Science (Biology), Associate Professor, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: samusenk@mail.ru 

Yuriev Anatoliy Leonidovich, Candidate of Science (Biology), Associate Professor, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: yuriev@bk.ru 

Vokin Aleksey Innokentyevich, Candidate of Science (Biology), Associate Professor, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: vokin@bk.ru 

Samusenok Innokentiy Vitalyevich, Leading Engineer, Irkutsk State University, 1, K. Marx st., Irkutsk, 664003, Russian Federation, e-mail: aen_seidhe@mail.ru 

Alekseyev Sergey Sergeevich, Doctor of Science (Biology), Senior Research Scientist, Koltzov Institute of Developmental Biology RAS, 26, Vavilov st., Moscow, 119991, Russian Federation; A.N. Severtsov Institute of Ecology and Evolution RAS, 33, Leninski av., Moscow, 119071, Russian Federation, e-mail: alekseyev@mail.ru

Andreev R.S., Matveev A.N., Samusenok V.P., Yuriev A.L., Vokin A.I., Samusenok I.V., Alekseyev S.S. Sculpin Cottus cf. poecilopus Heckel, 1837 in Baikal Lake Basin: First Findings. The Bulletin of Irkutsk State University. Series Biology. Ecology, 2020, vol. 31, pp. 30-51. https://doi.org/10.26516/2073-3372.2020.31.30 (in Russian)

ichthyofauna, Cottus cf. poecilopus, introduction, age structure, growth, feeding habits, Baikal Lake basin, Lena River basin

Berg L.S. Zametki po sistematike baikal'skikh Cottidae [Notes on the systematics of Baikalian Cottidae]. Ezhegodnik Zoologicheskogo muzeya Imperatorskoj Akademii Nauk [Ann. Zool. Museum Emper. AS], 1903, vol. 8, pp. 99-114. (in Russian)

Berg L.S. Ryby presnykh vod SSSR i sopredelnykh stran [Freshwater fishes of USSR and adjacent territories. Part 3]. Moscow, Saint Petersburg, AS USSR Publ., 1949, 1381 p. (in Russian)

Matveev A.N., Samusenok V.P., Pronin N.M., Tel'pukhovskii A.N., Vokin A.I., Prosekin K.A., Yur'ev A.L. Biologiya novogo podvida sibirskogo khariusa Thymallus arcticus baisalolenensis ssp. nova (Salmoniformes, Thymallidae) v basseine ozera Baikal [Biology of new subspecies of Siberian grayling Thymallus arcticus baisalolenensis ssp. nova (Salmoniformes, Thymallidae) in Baikal Lake basin]. Vestnik Buryatskogo gosudarstvennogo universiteta. Ser. biol. [Bull. Buryat St. Univ. Ser. Biol.], vol. 8, pp. 222-233. (in Russian)

Bogdanov B.E., Knizhin I.B. Osobennosti pitaniya pestronogogo podkamenshchika Cottus poecilopus (Cottisae, Scorpaeniformes) v vodoemakh razlichnogo tipa basseina Leny [Feeding habit of Cottus poecilopus (Cottisae, Scorpaeniformes) in different water bodies of Lena River basin]. Okhrana i nauchnye issledovaniya na osobo okhranyaemykh territoriyakh Dalnego Vostoka Sibiri [Protecting and researches in protected areas of Far East and Siberia. Proc. of the Intern. Sci. and Pract. Conf.]. Khabarovsk, 2007, pp. 19-27. (in Russian)

Bogdanov B.E., Sverdlova T.V., Knizhin I.B. Izmenchivost' i taksonomicheskii status sibirskikh populyatsii pestronogogo podkamenshchika Cottus poecilopus complex (Scorpaeniformes: Cottidae) [Variability and taxonomic status of siberian populations of Cottus poecilopus complex (Scorpaeniformes: Cottidae)]. Bull. Siber. Fed. Univ. Biology, 2013, vol. 6, no. 1, pp. 18-31. (in Russian)

Borisov P.G. Ryby reki Leny [Fishes of Lena River]. Proc. Yakut. Comm. AS USSR, 1928, vol. 9, pp. 1-188. (in Russian)

Vereshchagin G.Yu. Dva tipa biologicheskikh kompleksov Baikala [Two types of biological complexes in Baikal Lake]. Proc. Baikal Limnol. St., 1935, vol. 6, pp. 199-212. (in Russian)

Vokin A.I., Matveev A.N., Samusenok V.P., Yur'ev A.L. Vidovoe raznoobrazie i rasprostranenie khariusovykh ryb v vodoemakh Baikalo-Angarskogo basseina [Diversity and distribution of graylings in Baikal-Angara basin]. Sovremennye problemy biologii, ekologii i pochvovedeniya [Contemporary problems of biology, ecology and soil science: Proc. of the Intern. Sci. and Pract. Conf., Irkutsk, Russia]. Irkutsk, Irkutsk St. Univ. Publ., 2019, pp. 36-38. (in Russian)

Samusenok V.P., Alekseev S.S., Matveev A.N., Gordeeva N.V., Yur'ev A.L., Vokin A.I. Vtoraya v basseine Baikala i samaya vysokogornaya v Rossii populyatsiya arkticheskogo gol'tsa Salvelinus alpinus complex (Salmoniformes, Salmonidae) [Second in Baikal Lake basin and most alpine population of Arctic charr Salvelinus alpinus complex (Salmoniformes, Salmonidae)]. J. Ichthyol., 2006, vol. 46, no. 5, pp. 616-629. (in Russian)

Vyshegorodtsev A.A. Ryby Eniseya [Fishes of Yenisei River]. Novosibirsk, Nauka Publ., 2000, 237 p. (in Russian) 

Bazarov D.-D.B., Rezanov I.N., Budaev R.Ts., Imetkhenov A.B., Dergausova M.I., Rezanov V.P., Savinova V.V. Geomorfologiya Severnogo Pribaikaliya i Stanovogo nagor'ya [Geomorphology of Northern Cisbaikalia and Stanovoye upland]. Moscow, Nauka Publ., 1981, 198 p. (in Russian)

Golubtsov A.S., Malkov N.P. Ocherk ikhtiofauny Respubliki Altai: sistematicheskoe raznoobrazie, rasprostranenie i okhrana [Study of ichthyofauna of Altai Republic: systematic diversity, distribution and protection]. Moscow, KMK Publ., 2007, 164 p. (in Russian)

Karasev G.L. Ryby Zabaikaliya [Fishes of Transbaikalia]. Novosibirsk, Nauka Publ., 1987, 295 p. (in Russian)

Sideleva V.G., Neelov A.V., Voronina E.P., Volkova G.A. Katalog fondovoi kollektsii Zoologicheskogo instituta RAN. Klass Kostistye ryby (Osteichtyes). Otryad Skorpenoobraznye (Scorpaeniformes). Podotryad Cottoidei. Part 2: Issledovaniya fauny morei [Collection catalog of Institute of Zoology RAS. Osteichtyes. Scorpaeniformes: Cottoidei. Part 2: Researches of sea fauna], 2006, vol. 57, no. 65, pp. 225-349. (in Russian)

Kashchenko N.F. Rezultaty altaiskoi zoologicheskoi ekspeditsii 1898 g. Pozvonochnye [Results of Altay zoological expedition in 1898. Vertebrates]. Tomsk, Tomsk St. Univ. Publ., 1899, 158 p. (in Russian)

Kirillov F.N. Ryby Yakutii [Fishes of Yakutia]. Moscow, Nauka Publ., 1972, 360 p. (in Russian)

Knizhin I.B. Khariusy (Thymallus Cuvier, 1829) Golarktiki (sistematika, filogeografiya, osobennosti ekologii) [Graylings (Thymallus Cuvier, 1829) of Holarctycs (systematics, filogeography and ecology. Doctor in Biology dissertation abstract]. Moscow, Inst. Ecol. and Evol. RAS Publ., 2009, 267 p. (in Russian)

Kozhov M.M. Ozero Frolikha [Lake Frolikha]. Irkutsk, Irkutsk St. Publ., 1942, 32 p. (in Russian)

Kozhov M. M. Ocherki po baikalovedeniyu [Issues on Baikal Lake]. Irkutsk, Irkutsk St. Univ. Publ., 1972, 252 p. (in Russian)

Kononov E.E. O novykh dannykh po probleme Pra-Manzurskogo kanala stoka baikal'skikh vod [New data on the problem of the Pra-Manzurka runoff channel for Baikal waters]. Izvestiya SO RAEN. Geologiya, poiski i razvedka rudnykh mestorozhdenii [Rep. SB RAS. Geology, prospecting and exploration of ore deposits], 2016, no. 2, pp. 116-128. (in Russian). 

Koryakov E.A., Sidelev G.N. Bychki-podkamenshchiki iz ozera Agata (plato Putorana) [Sculpins of Agatha Lake (Putorana Plateau)]. J. Ichthyol., 1976, vol. 16, no. 3, pp. 553-555. (in Russian)

Kuz'min M.I. Vo ldakh Baikala [Amidst the Baikal ice]. Novosibirsk, Geo Publ., 2001, 140 p. (in Russian)

Lamakin V.V. Neotektonika Baikalskoi vpadiny [Neotectonics of Baikal hollow]. Moscow, Nauka Publ., 1968, 245 p. (in Russian)

Logachev N.A., Lomonosova T.K., Klimanova V.M. Kainozoiskie otlozheniya Irkutskogo amfiteatra [Cenozoic deposits of Irkutsk amphitheater]. Moscow, Nauka Publ., 1964, 195 p. (in Russian)

Logashev M.V. Ozero Melkoe i ego rybokhozyaistvennoe znachenie [Lake Melkoye and its fishery importance]. Tr. in-ta polyar. zemledeliya, zhivotnovodstva i promysl. khoz-va. Ser. «Promysl. khoz.-vo» [Proc. Inst. Polar Agriculture, Animal Husbandry and Hunting Industry and Fishery. Ser. Hunting Industry and Fishery]. 1941, vol. 11, pp. 7-71. (in Russian)

Matveev A.N. Struktura rybnogo naseleniya v vodoemakh Baikal'skoi riftovoi zony [Fish community structure in waterbodies of Baikal rift zone: Doctor in Biology dissertation abstract]. Irkutsk, Irkutsk St. Univ. Publ., 2006. 682 p. (in Russian)

Matveev A.N., Samusenok V.P. Bioraznoobrazie i struktura rybnogo naseleniya gornykh vodoemov Baikal'skoi riftovoi zony [Biodiversity and structure of fish communities in mountain water bodies of Baikal rift zone]. Byull. VSNTs SO RAMN [Bull. East-Sib. SC SB RAMS], 2006, no. 2, pp. 84-91. (in Russian)

Matveev A.N., Samusenok V.P. Proiskhozhdenie (istochniki i puti proniknoveniya) ikhtiofauny ozera Baikal [Origin of Lake Baikal ichthyofauna]. Razvitie zhizni v protsesse abioticheskikh izmenenii na Zemle [The process of life development within abiotic changes on the Earth: Sci. Conf., Listvyanka, Russia]. Novosibirsk, SB RAS Publ., 2008, pp. 401-414. (in Russian)

Mats V.D., Ufimtsev G.F., Mandel'baum M.M. Kainozoi Baikal'skoi riftovoi vpadiny: stroenie i geologicheskaya istoriya [The Cenozoic Baikal Rift Basin: Structure and Geological History]. Novosibirsk, SB RAS Publ., 2001, 252 p. (in Russian)

Mats V.D., Efimova I.M., Kul'chitskii A.A. Drevnie doliny zapadnogo Pribaikal'ya (istoriya formirovaniya) [Ancient valleys in the west Pribaikalye (history of formation)]. Geomorfologiya [Geomorfology], 2010, vol. 2, pp. 91-101. https://doi.org/10.15356/0435-4281-2010-2-91-101 (in Russian)

Mezhdunarodnyi kodeks zoologicheskoi nomenklatury. Izd. chetvertoe. Prinyat Mezhdunarodnym soyuzom biologicheskikh nauk [International codex of zoological nomenclature. 4^th ed.]. Moscow, KMK Publ., 2004, 223 p. (in Russian)

Metodicheskoe posobie po izucheniyu pitaniya i pishchevykh vzaimootnoshenii ryb v estestvennykh usloviyakh [Guide to study of feeding and feed relations in fishes in natural conditions]. Moscow, Nauka Publ., 1974, 254 p. (in Russian)

Mukhomed'yarov F.B. K biologo-sistematicheskoi kharakteristike [On biological characteristic and systematics of davatchan]. Proc. East-Siberian St. Univ., 1942, vol. 2, is. 3, pp. 119-126. (in Russian)

Florensov N.A. (ed.). Nagoriya Pribaikaliya i Zabaikaliya [Uplands of Cisbaikalia and Transbaikalia]. Moscow, Nauka Publ., 1974, 359 p. (in Russian)

Bogdanov B.E., Matveev A.N., Knizhin I.B., Samusenok V.P. Nekotorye dannye ob ekologo-ekologicheskikh osobennostyakh razmnozheniya trekh reofilnykh vidov podkamenshchikov (Cottidae) vodoemov Baikal'skogo regiona [Some data on the ecological and ecological characteristics of the reproduction of three rheophilic species of sculpin (Cottidae) in the water bodies of the Baikal region]. Tr. kaf. zool. pozv. [Proc. Chair Zool. Vert.]. Irkutsk, Irkutsk St. Univ. Publ., 2001, vol. 1, pp. 194-204. (in Russian)

Matveev A.N., Samusenok V.P., Tel'pukhovskii A.N., Pronin N.M., Vokin A.I., Prosekin K.A., Anoshko A.N. Novyi podvid sibirskogo khariusa Thymallus arcticus baisalolenensis ssp. nova (Salmoniformes, Thymallidae). Vestnik Buryatskogo gosydarstvennogo universiteta. Ser. biol. [Bull. Buryat St. Univ. Ser. Biol.], 2005, vol. 7, pp. 69-82. (in Russian)

Ol'shanskaya O.L. Obzor ikhtiofauny reki Pyasina [An overview of ichthyofauna of Pyasina River]. J. Ichthyol., 1965, vol. 5, no. 2, pp. 263-277. (in Russian)

Plokhinskii N.A. Biometriya [Biometrics]. Moscow, Nauka Publ., 1970, 368 p. (in Russian)

Pravdin I.F. Rukovodstvo po izucheniyu ryb (preimushchestvenno presnovodnykh) [Guide to studying of fishes]. Moscow, Pishch. prom-st Publ., 1966, 376 p. (in Russian)

Pavlov D.S., Savvaitova K.A., Gruzdeva M.A., Maksimov S.V., Mednikov B.M., Pichugin M.Yu., Savoskul S.P., Chebotareva Yu.V., Pavlov S.D. Raznoobrazie ryb Taimyra: Sistematika, ekologiya struktura vidov kak osnova bioraznoobraziya v vysokikh shirotakh, sovremennoe sostoyanie v usloviyakh antropogennogo vozdeistviya [Diversity of fishes of Taymyr Peninsula]. Moscow, Nauka Publ., 1999, 207 p. (in Russian)

Gordeeva N.V., Alekseev S.S., Kirillov A.F., Vokin A.I., Samusenok I.V. Rasprostranenie, sostav i rodstvennye otnosheniya filogeneticheskikh grupp arkticheskogo gol'tsa Salvelinus alpinus (Salmoniformes, Salmonidae) v evropeiskoi chasti Rossii i Sibiri po dannym analiza nukleotidnykh posledovatelnostei mitokhondrial'noi DNK [Distribution, composition and relationships of phylogenetic groups of Arctic charr Salvelinus alpinus (l.) (Salmoniformes, Salmonidae) in the european part of Russia and in Siberia as revealed by the analysis of nucleotide sequences of mitochondrial DNA]. J. Ichthyol., 2018, vol. 58, no. 6, pp. 659-669. (in Russian). http://doi.org/10.1134/S0042875218050107

Redkozubov I.Yu., Movchan V.A. K izucheniyu davatchana Salvelinus alpinus erythrinus (Georgi) oz. Frolikha [To study of davatchan Salvelinus alpinus erythrinus (Georgi) from Frolikha Lake]. J. Ichthyol., 1974, vol. 14, no. 2, pp. 330-332. (in Russian)

Pudovkina T.A., Sitnikova T.Ya., Matveev A.N., Shcherbakov D.Yu. Rodstvennye svyazi baikalskikh polikhet roda Manayunkia [Polychaeta: Sedentaria: Sabellidae] po dannym analiza CO1 s analizom istorii rasseleniya [The history of Manayunkia [Polychaeta: Sedentaria: Sabellidae] propagation in northeastern Asia]. Ekol. Gen., 2014, vol. 12, no. 3, pp. 32-42. (in Russian). https://doi.org/10.17816/ecogen12332-42

Romanov V.I. Ikhtiofauna Khantaiskoi gidrosistemy [Ichthyofauna of Khantaisky hydrosistem]. Priroda Khantaiskoi gidrosistemy [Nature of Khantaisky hydrosistem]. Tomsk, Tomsk St. Univ. Publ., 1988, pp. 199-236. (in Russian)

Pronin N.M., Matveev A.N., Samusenok V.P., Bobkov A.I., Sokolov A.V., Dzyumenko N.F., Kalyagin L.F., Gorlachev V.P., Pronina S.V., Dugarov Zh.N., Vokin A.I., Yur'ev A.L. Ryby ozera Baikal i ego basseina [Fish of lake Baikal and its basin]. Ulan-Ude, Buryat SC RAS Publ., 2007, 284 p. (in Russian)

Gundrizer A.N., Ioganzen B.G., Kafanova V.V., Krivoshchekov G.M. Ryby Teletskogo ozera [Fishes of Teletsky Lake]. Novosibirsk, Nauka Publ., 1981, 150 p. (in Russian)

Savvaitova K.A., Maksimov V.A., Medvedeva E.D. Davatchan Salvelinus alpinus erythrinus (Georgi). J. Icht., 1977, vol. 17, is. 2, pp. 203-219. (in Russian)

Samusenok V.P. Ekologiya arkticheskogo gol'tsa Salvelinus alpinus (L.) vysokogornykh vodoemov Severnogo Zabaikaliya [Ecology of Arctic charr Salvelinus alpinus (L.) in mountain water bodies of northern Transbaikalia: Candidate in Biology dissertation abstract]. Irkutsk, Irkutsk St. Univ. Publ., 2000. 214 p. (in Russian)

Sideleva V.G. Ryby [Fishes]. Annotirovannyi spisok fauny ozera Baikal i ego vodosbornogo basseina [Annotated list of Baikal Lake and Baikal basin fauna]. Timoshkin O.A., Sitnikova T.Ya., Rusinek O.T. (eds.). Novosibirsk, Nauka Publ., 2004, vol. 1: Baikal Lake, book 2, pp. 1024-1050. (in Russian)

Sideleva V.G., Goto A. Vidovoi status i pereopisanie trekh vidov gruppy Cottus poecilopus (Cottidae) Evrazii [Specific status and redescription of three species of group Cottus poecilopus (Cottidae) in Eurasia]. J. Ichthyol., 2009, vol. 49, no. 5, pp. 617-631. (in Russian)

Sideleva V.G., Goto A. Novyi vid podkamenshchika Cottus kolymensis sp. nova (Scorpaeniformes: Cottidae) iz rek Kolymskogo kraya [New species of sculpin Cottus kolymensis sp. nova (Scorpaeniformes: Cottidae) from the rivers of Kolyma Region]. J. Ichthyol., 2012, vol. 52, no. 3, pp. 288-294. (in Russian)

Sitnikova T.Ya., Prozorova L.A. Eshche raz o proiskhozhdenii baikal'skikh endemichnykh mollyuskov semeistva Baicaliidae Clessin 1880 (Gastropoda) [One more about origin of baikalian endemic molluscs Baicaliidae Clessin 1880 (Gastropoda)]. Razvitie zhizni v protsesse abioticheskikh izmenenii na Zemle [The process of life development within abiotic changes on the Earth: Sci. Conf., Listvyanka, Russia]. Novosibirsk, SB RAS Publ., 2008, pp. 371-375. (in Russian)

Taliev D.N. Bychki-podkamenshchiki Baikala [Sculpins of Baikal Lake (Cottoidei)]. Moscow, St.-Petersb., Nauka Publ., 1955, 603 p. (in Russian)

Taranets A.Ya. O nekotorykh novykh presnovodnykh rybakh iz Dalnevostochnogo kraya [On some new fresnwater fishes from Far East region]. Rep. AS USSR, 1933, no. 2, pp. 83-85. (in Russian)

Taranets A.Ya. Presnovodnye ryby basseina severo-zapadnoi chasti Yaponskogo morya [Freshwater fishes of basin of north-western part of Japan Sea]. Proc. Zool. Inst. AS USSR, 1936, vol. 4, is. 2, pp. 485-540. (in Russian)

Ufimtsev G.F., Shchetnikov A.A., Filinov I.A. Inversii v noveishei geodinamike Baikalskoi riftovoi zony [Neotectonic inversions in the Baikal rift system]. Russ. Geol. Geophys., 2009, vol. 50, no. 7, pp. 796-808. (in Russian)

Chereshnev I.A. Bychok-podkamenshchik roda Cottus (Cottidae) iz basseina r. Chaun (arkticheskaya Chukotka) [Sculpin Cottus (Cottidae) from Chaun River basin (arctic Chukcha Peninsula]. J. Ichthyol., 1982, vol. 22, no. 1, pp. 15-26. (in Russian)

Shed'ko S.V., Shed'ko M.B. Novye dannye po presnovodnoi ikhtiofaune yuga Dal'nego Vostoka Rossii [New data on freshwater fauna of southern Russian Far East]. V.Ya. Levanidov Memorial Scientific Readings. Vladivostok, Dalnauka Publ., 2003, vol. 2, pp. 319-336. (in Russian)

Shed'ko S.V., Miroshnichenko I.L. Filogeneticheskie svyazi podkamenshchika volka Cottus volki Taranetz, 1933 (Scorpaeniformes, Cottidae) soglasno rezultatam analiza kontrol'nogo uchastka mitokhondrialnoi dnk [Phylogenetic connections of Cottus volki Taranetz, 1933 (Scorpaeniformes, Cottidae) according to the analysis of the control region of mitochondrial DNA]. J. Ichthyol., 2007, vol. 47, no. 1, pp. 26-30. (in Russian). 

Bessudova A.Yu., Firsova A.D., Tomberg I.V., Sorokovikova L.M., Likhoshway Ye.V. Biodiversity of silica-scaled chrysophytes in tributaries of northern limit of Lake Baikal. Acta Biologica Sibirica, 2018, vol. 4, no. 3, pp. 75-84. http://dx.doi.org/10.14258/abs.v4i3.4411 

Chilton D.E., Beamich R.J. Age determination methods for fishes studied by the groundfish program of the Pacific Bological station. Can. Spec. Publ. Fish. Aquat. Sci. Ottawa, 1982, vol. 60, 102 p.

Dybowski B.N. Vorlaufige Muttheilungen uber die Fischfauna des Ononflusses and des Ingoda in Transbaicalien. Verh. Zool.-bot. Gez. Vien, 1869, Bd. 19, pp. 209-222. 

Freyhof J., Kottelat M., Nolte A. Taxonomic diversity of European Cottus with description of eight new species (Teleostei: Cottidae). Ichthyol. Explor. Freshwaters, 2005, vol. 16, no. 2, pp. 107-172.

Georgi J.G. Bemerkungen einer Reise im Russischen Reich im Jahre 1772. Bd. I, St. Petersb., Keiserl. Academie der Wissenschaften, 1775, 506 s. 

Kontula T., Väinölä R. Comparative phylogeography of freshwater cottid fishes: continent-scale versus European affinities in the Cottus poecilopus and Cottus gobio complex. Acad. dissertation. Helsinki, 2003, pp. 1-15.

Yokoyama R., Sideleva V.G., Shedko S.V., Goto A. Broad-scale phylogeography of the Palearctic freshwater fish Cottus poecilopus complex (Pisces: Cottidae). Mol. Phylogenet. Evol., 2008, vol. 48, pp. 1244-1251. http://doi.org/10.1016/j.ympev.2008.02.002